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The family Hominidae (great apes) diverged from the family Hylobatidae (gibbons) some 15–20 million years ago; subfamily Homininae (African apes) diverged from Ponginae (orangutans) about 14 million years ago; The tribe Hominini (including humans, Australopithecus, and chimpanzees) parted from the tribe Gorillini (gorillas) between 8–9 million years ago; and, in turn, the subtribes Hominina (humans and extinct biped ancestors) and Panina (chimpanzees) separated 4–7 million years ago. Anatomically modern humans appeared in Africa approximately 300,000 years ago.